The stark image of a solitary coyote is familiar to many, thanks to its use in countless Westerns. It has been a persistent image, and for long the coyote was considered to be a solitary animal. But recent studies have shown that in some situations coyotes live cooperatively in a way similar to wolves.

The coyote-whose name derives from the original Aztec word for the species, coyote-is a medium-sized canid with a rather narrow muzzle, large pointed ears and long slender legs. Size varies between populations and from one locale to another, and adult males are usually heavier and larger than adult females.
While the geographic ranges of most predators are shrinking, that of the coyote is increasing. A northerly and, particularly, an easterly expansion from the central Great Plains began in the late 19th century, as local populations of the larger canids, the Gray wolf (Canis lupus) and the Red wolf (C.rufus), were decimated by man.
Coyotes can interbreed with the Domestic dog, the Red wolf and, probably, the Gray wolf (the so-called Eastern coyotes are now thought to be fertile coyote-Gray wolf hybrids). The coyote-Domestic dog hybrid (“coydog”) can reproduce at one year old and has two litters a year. Coydogs are even more liable to attack farm and domestic animals than are coyotes.
Like jackals and wolves, the coyote is an opportunistic predator. Mammals, including carrion, generally make up over 90 percent of its diet. Ground squirrels, rabbits and mice predominate, but larger animals such as the Pronghorn antelope, deer and Rocky mountain sheep are included. Coyotes also eat fruit and insects. Small prey are hunted singly, but larger animals are hunted cooperatively. Coyotes normally stalk small prey from a few meters, but occasionally from as far as 50 meters (165 ft) and for as long as 15 minutes. Two or more coyotes may chase larger prey for up to 400m (1,300ft).
Both sexes attain sexual maturity during the first breeding season (January to March) following birth. Females produce one litter a year, averaging six pups per litter. The young are born blind and helpless in a den and are nursed for a period Of 5-7 weeks. At three weeks pups begin to eat semisolid food regurgitated by both parents and other pack members of both sexes. Most young disperse in their first year and may travel up to 160 km (100mi) before settling down.
The basic social unit in most coyote populations is the breeding pair; and the size of the home range varies from 14 to 65 sqkm (5.5 to 25 sq mi) for males, with an average Of 25 sqkm (9.9 sqmi) for females. Coyotes are now known to form packs similar to wolf-packs, in certain situations. Such packs are formed by delayed dispersal of the young, who remain as “helpers” in a pack; a typical pack consists of about six closely related adults, yearlings and young. It is usually the dominant male and female that breed.
Pack members sleep, travel and hunt larger prey together and cooperate in territorial disputes and defense of carrion. In general, coyote packs are smaller than wolf packs and associations between individuals less stable. The reasons for this may be the early expression of aggression, which is found in coyotes but not in wolves, and the fact that coyotes often mature in their first year whereas wolves do so in their second.
Variation in social organization enables the coyote to thrive on diverse prey and this flexibility is probably the reason for the wide, and expanding, geographic range of the species. Coyotes living in packs are more effective predators of large animals, and where such prey (eg deer, elk) is available, packs Of 3-8 coyotes are found. Where the principal prey is small mammals, the pups disperse early, packs are not formed, and most sightings are of solitary coyotes. Seasonal variation in social structure also occurs: when Ground squirrels and the young of large mammals are available as prey, coyotes spend less time together.
Coyotes use urine marking and calls to define their territory, to communicate with each other, and to strengthen social bonds. The coyote’s howl is unique and consists of a series of high-pitched staccato yelps followed by a prolonged siren wail. Their vocalizations also include barks, barkhowls, group yip-howls and group howls.
During the last 150 years, coyotes have been responsible for large economic losses to US agriculture, especially sheep farming. Some ranchers have lost up to 67 percent of their lambs and 20 percent of their sheep to coyotes in a single year; others lose very few. In fact, there is evidence that attempts to control coyotes by poisoning may also deplete the numbers of their natural prey and lead to increasing attacks by coyotes on farm animals. Although the species is not endangered, it is now totally protected in I2 states and the coyote harvest is regulated by a hunting or trapping season in most of the remaining states and Canada.


